17 18 19
Valley fold the diagonal Valley fold a parallel Crimp.
between the points of diagonal starting at
intersection and unfold. the intersection point 22
indicated. Rotate.
20
21
Reverse fold. Crimp. Valley fold the top flap.
23 24 25
Valley fold and unfold. Squash fold asymmetrically. Valley fold the sides into
Model will not lie flat. the center.
Luna Moth 101
26 27 28
Wrap layers behind. Valley fold. Valley fold while pulling
out some trapped paper.
29 30
31
Valley fold. Valley fold and unfold. Fold and unfold into
fourths.
32 33 34
22-31
Mirror steps 22 to 31 on the Valley fold. Valley fold while pulling
flap behind. out some trapped paper.
102 Origami Masters: Bugs
35 36 37
Valley fold. Open sink. Valley fold while raising
center flap.
38 39
40
Valley fold the flap up. Valley fold and unfold. Fold and unfold in fourths.
41 42 43
Valley fold. Pleat. Unfold.
Luna Moth 103
44 45 46
Valley fold edge to crease Valley fold. Valley fold the diagonal
mark indicated. connecting indicated
intersection points.
47 48
49
Valley fold, aligning the Valley fold edge to Unfold.
folded edge with the crease folded edge.
mark indicated. 52
50 51
Reverse fold. Crimp. Valley fold through points
104 Origami Masters: Bugs of intersection.
53 54 55
Open unsink. Open unsink. Spread sink.
56 57 58
Spread sink. Closer view. Valley fold and unfold.
59 60 61
Petal fold. Unfold. Open sink.
Luna Moth 105
62 63 64
Larger view. Turn over. Spread sink.
65 66 67
Spread sink. Closed unsink. Squash fold along existing
creases.
68 69
70
Valley fold down. Collapse into a waterbomb Squash fold while lifting the
106 Origami Masters: Bugs base by pressing in the sides. bottom center edge.
71 72 73
Turn over. Squash fold. Open sink in and out.
74 75 76
Squash fold. Spread sink the top while Valley fold the sides to
squash folding. The model the center.
77 will not lie flat.
78 79
Wrap layers behind. Valley fold. Valley fold while pulling
out some trapped paper.
The left side will not lie flat.
Luna Moth 107
80 81 82
74–81
Valley fold. Open sink in and out. Mirror steps 74 to 81
on the flap behind.
83
84 85
Valley fold. The left side Open sink in and out. Valley fold.
will not lie flat.
88
86 87
Spread sink. The model Closed unsink. Valley fold down.
will not lie flat.
108 Origami Masters: Bugs
89 90 91
Valley fold. 86–89
92 Mirror steps 86 to 89 on the Closer view.
flap behind.
93 94
Valley fold. Pleat. Valley fold.
95 96 97
Valley fold down. Valley fold and unfold. Fold and unfold in fourths.
Luna Moth 109
98 99 100
Unfold. Crimp. Petal fold.
101 102 103
Valley fold down. Open sink. Spread sink while lifting
the center flap.
104 105
106
Closed unsink. Valley fold up. Spread sink.
110 Origami Masters: Bugs
107 108 Closed 109
Valley fold. Open sink, except for Crimp.
the top corner, which
110 will be closed. 112
111
Open sink in and out. Valley fold. Valley fold and unfold.
113 114 115
92–114
Open sink. Valley fold. Mirror steps 92 to 114 on
the flap behind.
Luna Moth 111
116 117 118
Valley fold while squash Valley fold down. Valley fold and unfold.
folding interior flap.
121
119 120
Open sink. Valley fold up. Turn over.
122 123 124
Valley fold. Closed unsink. Valley fold and unfold
112 Origami Masters: Bugs the corners.
125 126 127
Reverse fold. Valley fold and unfold Valley fold and unfold
the corners. in fourths.
128
129 130
Open sink in and out on Valley fold and unfold. Pleat while pushing out
both corners. paper trapped inside.
131 132 133
Valley fold up. Closer view. Squash fold.
Luna Moth 113
134 135 136
Valley fold. Open sink. Squash fold.
137 138 139
Valley fold down. Pleat while pushing In progress.
interior layer in.
140 141 142
Pleat while collapsing In progress. Open paper and swivel fold.
paper inside.
114 Origami Masters: Bugs
143 144 145
In progress. Reverse fold. Reverse fold.
146 147 148
Reverse fold. Valley fold and unfold. Hybrid sink (top is closed,
bottom is open).
149
150 151 132–149
Larger view. Valley fold up. Repeat steps 132 to 149
on the other side of the
same flap.
Luna Moth 115
152 153 132–152 154
Valley fold. Mirror steps 132 to 152 Turn over.
on the other side.
155
156 157
Swivel fold inside. Model Rabbit-ear. Fold sides down while
will not lie flat. turning the long flap to
give it dimension.
158 159
160
Mountain fold the center 155–158 Closer view.
portion of the long flap.
116 Origami Masters: Bugs Mirror steps 155 to 158 on
the other side.
161 162 163
Valley fold down while Valley fold down. Swivel fold inside.
folding the sides inside.
166
164 165
Squash fold inside. Squash fold inside. Squash fold inside.
167 168 169
163–166
Mirror steps 163 to 166 on Turn over. Petal fold on both sides.
the other side. Luna Moth 117
170 171 172
Valley fold the corners. Rabbit-ear fold. Pleat.
173 174 175
Valley fold up. Valley fold up. Swivel bottom two flaps to
the right while folding the
176 177 top flap down. This will free
some paper on the left.
178
Spread sink. The model Closed unsink. Valley fold down.
will not lie flat.
118 Origami Masters: Bugs
179 180 181
Valley fold over. Valley fold up. Squash fold.
182 183 184
Kite fold in thirds. Valley fold up. Squash fold.
185 186 187
Kite fold. Kite fold in thirds. Valley fold up.
Luna Moth 119
188 189 190
Squash fold. Kite fold. Kite fold in thirds.
191 192 193
Valley fold down. Valley fold up. Valley fold over.
194 195 196
172–192 Squash fold. Reverse fold on both sides.
Mirror steps 172 to 192
on the other side.
120 Origami Masters: Bugs
197 198
Open antennae Mountain fold the
and spread. Model model in half. Rotate.
will not lie flat.
199
Valley fold a leg down.
200
Valley fold to make the leg jointed. Folded luna moth set against the crease pattern
for this model.
201
Luna Moth 121
199–200
Repeat steps 199 to 200 five more
times to shape remaining legs.
SHuKI KATo
Flying Hercules
Beetle
Titan Beetle Shuki Kato first learned about origami from
John Montroll’s classic book, Origami
122 Origami Masters: Bug Wars for the Enthusiast, at around age 6.
Shortly afterward, he began designing
his own original models. In
2000 he became a member of
OrigamiUSA and in college he
started giving lectures about origami
design as well as teaching classes on simple
origami models and folding techniques.
An avid golfer, he also enjoys studying about
animals (primarily dinosaurs and insects) and is
a fan of fantasy novels and Japanese manga and
anime. Shuki Kato currently resides in Kalispell,
Montana, and is working on a comprehensive
origami book of his own.
Photographs, crease patterns, and descriptions of
much of his work can be found at his
Flickr account:
www.flickr.com/photos/origami-artist-galen/
TITAn 1
BeeTLe
I first designed a version of this model in 2008 and revised it in 2012.
I like to use tracing paper to fold this model. I recommend starting out
a finished piece of about 6¼ inches.
with a 25-inch square, which results in
It takes me 7 hours to fold this particular model.
Start with the colored side
wiythWoestuorhhemweetnwoehewrehvkhomeoarlenotI,dmofaeononlledddstesiwtlelhlcaietlttihnieoostncnrofcalamaect,tip,natlhIgetestipenmparttepahh.yeeeIrft,fphiyInaeouapumsleersaotapwdeppewillslyl.ewltTtiiokfhtiohenisliymwsphsaura.otcgeAchreat,swnfstlhadawettewotlrerarnkosintsrakpbsnlopeedsir.ntadtiyrfy up. Fold in half vertically
and unfold.
2 34
Fold and unfold, creasing Fold and unfold, creasing Fold and unfold horizontally
only where shown. only where shown. and vertically through the
intersection of the creases.
5 67
Fold and unfold horizontally Fold the diagonal creases Fold and unfold the outer
and vertically to the creases and unfold. boxes in the center row.
indicated. This will divide Do not crease through
the square into three equal the center box.
rows of three squares.
Titan Beetle 123
8 9 10
Fold the corners to the Add four creases, creasing Fold and unfold through
center crease, pinch, and only where they intersect the crease intersections.
unfold. the diagonals.
11 12 13
Fold and unfold the side Fold creases to creases Fold edges to creases
edges to the creases as and unfold. shown and unfold.
shown.
14 15 16
Fold the bottom edge to the Fold the top edge to the Fold creases to creases as
crease indicated and unfold. crease indicated, crease the shown and unfold.
sides only, and unfold.
124 Origami Masters: Bugs
17 18 19
Fold edges to creases shown You should now have an Make additional horizontal
and unfold. incomplete 15 x 15 grid. and vertical creases shown,
folding creases to creases,
and unfold.
20 21
Fold the diagonals between the points indicated Fold the 18 angle bisectors shown and unfold.
and unfold. Turn over.
Titan Beetle 125
22 23
Fold the 16 angle bisectors shown and unfold. Divide rows shown in half horizontally, folding
Turn over. edge to crease or crease to crease, only where
indicated.
24
25
Divide more areas in half vertically and diagonally, Continue dividing more rows in half horizontally,
creasing between points indicated. Turn over. creasing between points indicated.
126 Origami Masters: Bugs
26 27
Continue dividing, adding vertical and diagonal Focus on this area.
creases between the points indicated.
30
28 29
Crease as shown, forming a number Pleat along existing creases. Pleat to the center along
of small diamonds. Turn over. existing creases.
31 32 33
Push the sides in to form a triangle Close up. Open sink the corners. Wrap one layer in front
in the center as you pleat down. on both sides.
Titan Beetle 127
34 35 36
Fold the two hidden Pull out the center region Unsink the sides.
edges down. as you pleat down.
37 38 39
Closed sink the edge. Pleat downward as you In progress.
collapse the center on
40 existing creases. 42
41
Bring one layer to the front Slightly larger view. Pleat to the center on
(closed sink). existing creases.
128 Origami Masters: Bugs
43 44 45 43–44
Sink the edge in and out on The gray region shows the Mirror steps 43 to 44
existing creases. result of the previous step. on the opposite side.
Open sink the top edge.
48
46 47
Turn the model over. Sink the hidden edge upward. Fold the edge down.
49 5047–48
Repeat steps 47 to 48 on the Pleat to the center, then
other three hidden edges. return to full view.
Titan Beetle 129
51 52
Collapse on existing creases. The This is what the collapse looks like on the
solid lines (mostly diagonals) indicate other side.
alternating valley and mountain folds.
Note: this is not for the faint of heart!
53 54 55
This diagram shows the Mandibles Antennae
folds that are obscured by Mouthparts (palps) Forelegs
four sets of pleats. Middle
130 Origami Masters: Bugs Head legs
Hind
Prothorax legs
Scutellum
Abdomen
Elytra
Extra
flap
In progress. This shows It should look like this. Note
only the most complicated the position of each flap.
part of the collapse, but it
will help orient you.
56 57 58
Spread the layers at Box-pleat the top edge Turn over from top to
the top apart. inside. bottom.
59 60 61
Fold the extra flap down. Pleat upward. You will have Side view of the
to readjust the diagonal folds previous step.
62 inside to accomplish this.
64
63
Position this way. Return to Pleat upward in the same Pleat upward in the same
view shown in step 60. manner as before. manner one more time.
Titan Beetle 131
65 66 67
Fold the thick abdomen Fold one layer over, squash Partial view, close up. Pull
down. folding the corner. The out some loose paper
model will not lie flat. (unsink).
68 69 70
Swivel the edge. Fold the edge over as Close back up.
indicated.
71
72 73
Fold to the center as you Wrap behind. Swivel this bit of paper
squash fold the top corner. inside.
132 Origami Masters: Bugs
74 75 76
66–73
Mirror steps 66 to 73 on the Fold the abdomen back up. Pleat in two places.
opposite side.
77 78 79
Unfold the last three pleats. Fold the sides behind on Swivel the small corners
existing creases. behind.
80 81 82
180˚
Reform the pleats, wrapping Mountain fold the small Position like this. Rotate
them all the way around hidden corners. 180º and return to full view.
the sides.
Titan Beetle 133
83 84 85
Fold the first pair of legs Pleat the next pair of legs Pleat the final pair of legs.
out to the sides. straight out.
86 87 88
Closer view. Fold one flap down. Fold and unfold.
89 90 91
Fold to the crease line. Fold the top layer inside. Fold over along
134 Origami Masters: Bugs the raw edge.
92 93 94
Rabbit-ear the small corner. Noting the reference, fold Fold in half, squash folding
the small flap back up. the lower corner.
95 96 97
Bring two layers Swivel the flap to the left. Squeeze the tiny flap in
to the front. half.
98 9987–97 100
Mirror steps 87 to 97 on It should look like this. Focus on the hind leg.
the opposite side. Return to full view. Titan Beetle 135
101 102 103
Lift the top layer. Fold the corner over and Fold the layer back.
over.
104
105 101–104 106
Position this way. Return to Mirror steps 101 to 104 on Turn the model over.
full view. the other hind leg.
107 108 109
Focus on the head. Squash fold the eyes. Wrap the layers behind.
136 Origami Masters: Bugs
110 111 112
Fold the small corners to Fold the top edge down Unsink one layer upward.
the sides. while pulling some paper
out at the corners.
113
114 115
Pull out some more Reverse fold the tiny Fold the eyes back toward
loose paper. corners and fold the top the center.
edge inside.
116 118
117
Fold the bottom edges Position this way. Return Focus on a hind leg.
behind and pull out a bit of to full view.
paper to shape the eyes. 121
119 120
Spread sink the tip. Fold the small flap to the Push a hidden edge inside as you
right. pleat one layer to the left.
Titan Beetle 137
122.
as you sq
122 123 124
Push a hidden edge in as Fold the small flap Push a hidden edge in and
you squash the small flap. to the right. pleat to the right.
125 126 127
Fold the small flap back to Make a tiny pleat. Fold the entire leg in half.
the right.
128 129 130
Fold and unfold. Reverse fold the top edge. Reverse fold.
hidden edge
heidsdmeanllefdlagpe.
e small flap.
en edge 131 132 133
mall flap.
Reverse fold. Narrow the end of the leg Crimp upward.
138 Origami Masters: Bugs and spread out the claws.
134 135 136
119–134 119–134
It should look like this.
Return to full view.
Repeat steps 119 to 134 on Crimp all the legs.
the remaining legs.
137 138 139
Sink two hidden Shape the wing Mountain fold the antennae
edges upward. covers (elytra). out to the sides.
140 141 142
Narrow the antennae. Crimp the antennae. Pleat the antennae, curve
the mandibles, and crimp
the mouthparts.
Titan Beetle 139
143 144
Narrow the antennae Continue shaping the model
further and give the ends to taste.
a bit of curve.
Folded titan beetle set against
crease pattern for this model
140 Origami Masters: Bugs
1
FLYIng Start with the colored side
HeRcuLeS up. Fold in half diagonally
and unfold.
BeeTLe
This is a recent (2012) revision of a model I first created in 2005. I folded
the model you see here with O-gami, but basically I would advise using the
thinnest paper available. Tracing paper is 41 gsm, so try anything thinner
than 50 gsm. To get a finished model with a 6½-inch wingspan, I started
with a 17-inch square. Leave yourself at least 5 to 7 hours to fold this model.
23 4
Fold in half vertically, Fold the corner to the Fold the top corner to the
making a pinch at the top. crease, pinch the left edge bottom edge, creasing
and unfold. lightly through the pinch
made in step 3, and unfold.
56
7
Fold the bottom corner Fold to the pinch Fold and unfold. This will
to the crease intersection, mark and unfold. create a rectangle.
pinch, and unfold.
Flying Hercules Beetle 141
8 9 10
Divide the rectangle into Fold the diagonal creases. Fold the last fourth in half,
fourths. Note that the creasing as indicated.
creases from steps 2 to 5
are no longer shown.
11 12 13
Valley fold. Check the next The points indicated should Fold, aligning points as
step for references. line up as shown. indicated, and unfold.
14 15 16
Fold the edge down; note that Close up view shows the Fold back down.
the reference is not exact and approximate reference is at
lies slightly above where you the ¹₅ mark.
would expect it to be.
142 Origami Masters: Bugs
17 18 19
Fold between the points Fold along the angle Fold and unfold.
indicated and unfold. bisector and unfold.
22
11–19 21
20
The model should look like Turn the paper over. Fold the 16 angle bisectors
this. Mirror steps 11 to 19 shown and unfold.
on the opposite side.
25
23 24
45°
Fold the 30 angle bisectors Fold in half diagonally. Reverse fold the corners along
shown and unfold. Rotate counterclockwise 45º. the top edge. From this point to
step 36 you won’t need to crease
firmly unless there is an already
existing crease.
Flying Hercules Beetle 143
26 27 28
Pleat to the left. Repeat Petal fold, swinging the Squash the layer to the
behind. excess paper to the left. right.
The model will not lie flat.
29
30 31
Squash the top edge Rabbit-ear fold as shown. Fold two flaps down.
and petal fold.
32 33 34
Fold one flap back up. Note that the creases should line Fold the flap back down.
144 Origami Masters: Bugs up (due to the bizarre reference
in step 15). The crease on the left
would be higher if you folded
the edge too far down.
35 27–34 36 37
Repeat steps 27 to 34 Fold the layer behind to the Open up along the center.
behind. right. The next steps will focus on
this area only.
38
39 40
Invert the pocket and Fold and unfold, noting Fold and unfold. This
swing the excess paper the references. should divide the area
to the right. into equal thirds.
41 42 43
39–40
Fold the middle flap to the Mirror steps 39 to 40 on the Collapse as shown. It is
left. opposite side. similar to a petal fold.
Flying Hercules Beetle 145
44 45 46
Fold corner to corner Fold the flap into Divide further.
and unfold. fourths and unfold.
49
47 48
Divide further. Make two outside crimps Squash fold.
on existing creases.
50
51 52
Petal fold, creasing the Fold and unfold. Swivel the top layer slightly.
mountain folds lightly.
146 Origami Masters: Bugs
53 54 51–53 55
Wrap to the inside and Mirror steps 51 to 53 on the Bring the triangle to
flatten. opposite side. the front (closed sink
it from behind).
56 57 58
Fold the corner and It should look like this. Turn the model over.
tuck it inside. Return to full view.
61
59 60
Petal fold, swinging the Invert the pocket (unsink). Swivel the top layers to the
excess paper to the left. right while pleating a hidden
layer.
Flying Hercules Beetle 147
62 63 64
Bring the trapped edge to Fold and unfold. Divide into fourths.
the front (unsink).
67
65 66
Fold one layer to the left. Fold and unfold, using the Fold one layer to the left.
creases behind as guides.
68 69 70
Fold and unfold, using the Fold the flap down. Divide into fourths.
creases behind as guides.
148 Origami Masters: Bugs
71 72 7367–68
Fold one layer to the left. Repeat steps 67 and 68 on Fold all the layers to
the next three layers. the right.
74 75 7660–73
Fold the large flap to the Repeat steps 60 to 73 on Squash fold the large flap.
right. this side.
77 78 79
Fold up as far as possible. Fold the corner and Squash fold the flaps down.
tuck it inside. Flying Hercules Beetle 149
80 81 82
Fold the flap down just past Swivel the edges out. Bring one layer to the front.
the straight edge.
83 84 85
Fold one layer to the left. Fold to the center. If the Fold inner edge to the
crease doesn’t touch the center.
point indicated, it isn’t
that important. 88
86 87
Bring the edge out Petal fold the edge up at It should look like this. Fold
from behind. an angle. Check the next the edge back inside.
150 Origami Masters: Bugs step for references.